M_1 Glucose_G6P ADPGK G6PC G6PC2 G6PC3 GALM GCK HK1 HK2 HK3 HKDC1 M_2 G6P_G3P ALDOA ALDOB ALDOC FBP1 FBP2 GPI PFKL PFKM PFKP TPI1 M_3 G3P_3PD BPGM COQ2 GAPDH GAPDHS PGAM1 PGAM2 PGAM4 PGK1 PGK2 PHB PPT1 M_4 3PD_Pyruvate BPGM COQ2 ENO1 ENO2 ENO3 MINPP1 PCK1 PCK2 PGAM1 PGAM2 PGAM4 PHB PKLR PKM PPT1 M_5 Pyruvate_Acetyl-Coa DLAT DLD PDHA1 PDHA2 PDHB M_6 Pyruvate_Lactate LDHA LDHAL6A LDHAL6B LDHB LDHC LDHD M_7 Acetyl-CoA+Oxaloacetate_Citrate ACLY CS M_8 Citrate_2OG ACO1 ACO2 IDH1 IDH2 IDH3A IDH3B IDH3G M_9 2OG_Succinyl-CoA DLD DLST OGDH OGDHL M_10 Succinyl-CoA_Succinate SUCLA2 SUCLG1 SUCLG2 M_11 Succinate_Fumarate SDHA SDHB M_12 Fumarate_Malate FH M_13 Malate_Oxaloacetate MDH1 MDH2 M_14 Pyruvate_Oxaloacetate PC M_15 3PD_Serine PHGDH PSAT1 PSPH M_16 Serine_Pyruvate SDS SDSL M_17 Serine_Cysteine CBS CTH M_18 Serine_Glycine AGXT SHMT1 SHMT2 M_19 Glycine_Creatine GAMT GATM M_20 Glycine_Glyoxylate AGXT AGXT2 DAO M_21 Choline_Glycine ALDH7A1 BHMT CHDH DMGDH PIPOX SARDH M_22 Glycine_Glycine-OUT ALAS1 ALAS2 AOC2 AOC3 GCAT GLDC MAOA MAOB M_23 Glycine_Glutathione GSS M_24 Glutathione_Glycine + Cysteine ANPEP GCLC GGT6 GSS LAP3 M_25 Glutathione_glutamate GGCT GGT1 GGT5 GGT6 GGT7 GSTA1 GSTA2 GSTA3 GSTA4 GSTA5 GSTK1 GSTM1 GSTM2 GSTM3 GSTM4 GSTM5 GSTO1 GSTO2 GSTP1 GSTT2 GSTT2B MGST1 MGST2 MGST3 OPLAH M_26 Glutamate + Cysteine_Glutathione GCLC GSS M_27 Serine_2-Aminoacrylate SDS SDSL M_28 Serine_Methionine BHMT BHMT2 CBS KYAT1 KYAT3 MTR M_29 Cysteine_Pyruvate CDO1 CTH GOT1 GOT2 KYAT1 KYAT3 MPST TST M_30 Methionine_Cysteine AHCY AHCYL1 AHCYL2 CBS CTH DNMT1 DNMT3A DNMT3B DNMT3L MAT1A MAT2A MAT2B M_31 Methionine_Methionine-OUT ADI1 AMD1 APIP ENOPH1 IL4I1 MAT1A MAT2A MAT2B MRI1 MTAP SMS SRM TAT M_32 Threonine_Propanoyl-CoA BCKDHA BCKDHB DBT DLAT DLD SDS SDSL M_33 G3P_PRPP ALDOA ALDOB ALDOC FBP1 FBP2 G6PD GPI H6PD PFKL PFKM PFKP PGD PGLS PGM1 PGM2 PRPS1 PRPS1L1 PRPS2 RBKS RPE RPEL1 RPIA TALDO1 TKT TKTL1 TKTL2 M_34 Acetyl-CoA_Fatty Acid ACACA ACACB ACSBG1 ACSBG2 ACSL1 ACSL3 ACSL4 ACSL5 ACSL6 FASN HSD11B2 HSD17B1 HSD17B12 HSD17B2 HSD17B6 HSD17B7 HSD17B8 MAST1 MCAT MECR OLAH OXSM PECR RDH10 RDH16 RDH8 RNF2 SLC27A2 M_35 Fatty Acid_Acetyl-CoA ACAA1 ACAA2 ACADL ACADM ACADS ACADSB ACADVL ACAT1 ACAT2 ACOX1 ACOX3 ACSBG1 ACSBG2 ACSL1 ACSL3 ACSL4 ACSL5 ACSL6 CPT1A CPT1B CPT1C CPT2 ECHS1 EHHADH GCDH HADH HADHA HADHB HSD17B10 HSD17B4 SLC27A2 M_36 Aspartate_Pyrimidine CAD DHODH M_37 Aspartate_Aspartate-OUT ACY3 ASPA FOLH1 NAALAD2 NAALADL1 NAT8L RIMKLA M_38 Aspartate_Asparagine ALPP ASNS ASPG ASRGL1 M_39 Aspartate_Fumarate ADSL ADSS ADSSL1 ASL ASS1 M_40 Aspartate_Oxaloacetate GOT1 GOT2 IL4I1 M_41 Aspartate_B-Alanine AZIN2 GAD1 GAD2 GADL1 M_42 Spermine_B-Alanine ALDH1A3 ALDH1B1 ALDH2 ALDH3A1 ALDH3A2 ALDH3B1 ALDH3B2 ALDH7A1 ALDH9A1 AOC2 AOC3 SMOX M_43 Histidine_B-Alanine CARNS1 CNDP1 CNDP2 M_44 B-Alanine_Acetyl-CoA ABAT ALDH6A1 M_45 Pyrimidine_B-Alanine DPYD DPYS NT5C NT5C1A NT5C1B NT5C1B-RDH14 NT5C2 NT5C3A NT5C3B NT5E NT5M PNP TK1 TK2 TYMP UCK1 UCK2 UCKL1 UPB1 UPP1 UPP2 UPRT M_46 Propanoyl-CoA_Succinate ECHDC1 MCEE MUT PCCA PCCB SUCLA2 SUCLG1 SUCLG2 M_47 Propanoyl-CoA_Acetyl-CoA ACACA ACACB ACADS ACOX1 ACOX3 ECHS1 EHHADH HADHA MLYCD M_48 Glutamate_Glutamine GLS GLS2 GLUL M_49 Glutamate_GABA GAD1 GAD2 M_50 GABA_Succinate ABAT ALDH5A1 M_51 Glutamate_2OG GLUD1 GLUD2 M_52 Histidine_glutamate AMDHD1 FTCD HAL UROC1 M_53 Leucine_Acetyl-CoA AACS ACADM ACAT1 ACAT2 AUH BCAT1 BCAT2 BCKDHA BCKDHB DBT DLD ECHS1 EHHADH HADHA HMGCL HMGCS1 HMGCS2 IVD MCCC1 MCCC2 OXCT1 OXCT2 M_54 Valine_Succinyl-CoA ABAT ACAD8 ACADM ACADS ACADSB ACSF3 AGXT2 ALDH1B1 ALDH2 ALDH3A2 ALDH7A1 ALDH9A1 AOX1 BCAT1 BCAT2 BCKDHA BCKDHB DBT DLD ECHS1 EHHADH HADH HADHA HIBADH HIBCH HSD17B10 MUT M_55 Isoleucine_Succinyl-CoA ACAA1 ACAA2 ACADM ACADS ACADSB BCAT1 BCAT2 BCKDHA BCKDHB DBT DLD ECHS1 EHHADH HADH HADHA HADHB HSD17B10 IL4I1 MCEE MUT PCCA PCCB M_56 Isoleucine_Acetyl-CoA ACAA1 ACAA2 ACADM ACADS ACADSB BCAT1 BCAT2 BCKDHA BCKDHB DBT DLD ECHS1 EHHADH HADH HADHA HADHB HSD17B10 IL4I1 M_57 Tyrosine_Fumarate FAH GOT1 GOT2 GSTZ1 HGD HPD IL4I1 MIF TAT M_58 Tyrosine_Tyrosine-OUT ALDH1A3 ALDH3A1 ALDH3B1 ALDH3B2 AOC2 AOC3 DDC MAOA MAOB TH TPO TYR M_59 Phenylalanine_Tyrosine PAH M_60 lysine_Acetyl-CoA AADAT AASS ACAT1 ACAT2 ALDH7A1 DLD ECHS1 EHHADH GCDH HADH HADHA HSD17B10 HSD17B4 HYKK PHYKPL M_61 Putrescine_GABA ALDH1B1 ALDH2 ALDH3A2 ALDH7A1 ALDH9A1 AOC1 M_62 Ornithine_Putrescine ODC1 M_63 Arginine_Ornithine ARG1 ARG2 M_64 Arginine_Putrescine AGMAT AZIN2 M_65 Argininosuccinate_Arginine + Fumarate ASL M_66 Citruline + Aspartate_Argininosuccinate ASS1 M_67 Ornithine_Proline OAT PRODH PRODH2 PYCR1 PYCR2 PYCR3 M_68 Proline_Glyoxylate + pyruvate ALDH4A1 GOT1 GOT2 HOGA1 PRODH2 PYCR1 PYCR2 PYCR3 M_69 Putresine_Spermine SMOX SMS SRM M_70 Methionine_Spermine AMD1 MAT1A MAT2A MAT2B SMOX SMS SRM M_71 Glucose_in_Glucose SLC2A1 SLC2A2 SLC2A3 SLC2A4 M_72 Acetyl-CoA_in_Acetyl-CoA SLC33A1 M_73 Citrate_in_Citrate SLC13A5 SLC25A1 M_74 2OG_in_2OG SLC25A11 M_75 Succinate_in_Succinate SLC13A3 SLC25A10 M_76 Malate_in_Malate SLC25A10 SLC25A11 M_77 Oxaloacetate_in_Oxaloacetate SLC25A21 M_78 Lactate_Lactate_out SLC16A1 SLC16A3 SLC16A4 M_79 serine_in_serine SLC1A4 SLC1A5 SLC38A2 SLC38A4 SLC38A5 SLC7A10 M_80 Cysteine_in_Cysteine SLC1A4 SLC1A5 SLC38A1 SLC38A2 SLC38A4 SLC7A10 SLC7A11 SLC7A7 M_81 Glycine_in_Glycine SLC6A5 SLC6A9 M_82 Creatine_in_Creatine M_83 Choline_in_Choline SLC5A7 M_84 Threonine_in_Threonine SLC1A4 SLC7A8 M_85 Aspartate_in_Aspartate SLC1A3 SLC1A6 M_86 Asparagine_in_Asparagine M_87 Pyrimidine_in_Pyrimidine SLC29A1 SLC29A2 M_88 Histidine_in_Histidine SLC15A4 M_89 B-Alanine_in_B-Alanine SLC6A6 M_90 Propanoyl-CoA_in_Propanoyl-CoA SLC27A1 M_91 glutamate_in_glutamate SLC17A6 SLC17A7 SLC17A8 SLC1A1 SLC1A2 SLC1A3 SLC1A5 SLC1A6 SLC1A7 M_92 Glutamine_in_Glutamine SLC1A5 M_93 GABA_in_GABA SLC32A1 SLC6A1 SLC6A11 SLC6A12 SLC6A13 M_94 Tyrosine_in_Tyrosine SLC7A5 SLC7A8 M_95 phenylalanine_in_phenylalanine SLC7A8 M_96 Valine_in_Valine SLC6A15 M_97 Isoleucine_in_Isoleucine SLC6A15 M_98 Leucine_in_Leucine SLC6A15 SLC7A5 SLC7A6 M_99 Lysine_in_Lysine SLC25A29 SLC38A4 SLC66A1 M_100 Arginine_Arginine SLC25A29 SLC7A1 SLC7A2 M_101 Citrulline_in_Citrulline SLC25A15 M_102 Ornithine_in_Ornithine SLC25A15 SLC25A15P1 SLC25A2 SLC25A29 M_103 Proline_in_Proline SLC36A1 SLC6A20 SLC6A7 M_104 Tryptophan_in_Tryptophan M_105 Fatty Acid_in_Fatty Acid SLC27A1 SLC27A2 SLC27A3 SLC27A4 SLC27A5 SLC27A6 M_106 Glucose_Glucose-6-phosphate GCK HK1 HK2 HK3 HKDC1 M_107 Glucose-6-phosphate_UDP-N-acetylglucosamine AMDHD2 GFPT1 GFPT2 GNPDA1 GNPDA2 GNPNAT1 GPI PGM3 UAP1 UAP1L1 M_108 Glucose-6-phosphate_Glucose-1-phosphate PGM1 PGM2 M_109 Glucose-1-phosphate_UDP-glucuronic acid UGDH UGP2 M_110 UDP-glucuronic acid+UDP-N-acetylglucosamine_Hyaluronic acid HAS1 HAS2 HAS3 M_111 Glucose-1-phosphate_Glycogen GYS1 GYS2 PYGB M_112 UDP-N-acetylglucosamine_CMP-N-acetylneuraminate CMAS GNE NANP NANS TSPAN31 M_113 Acetyl-CoA_(E,E)-Farnesyl-PP ACAT1 ACAT2 FDPS GGPS1 HMGCR HMGCS1 HMGCS2 IDI1 IDI2 MVD MVK PMVK M_114 (E,E)-Farnesyl-PP_Geranylgeranyl-PP FDPS GGPS1 M_115 (E,E)-Farnesyl-PP_Farnesal FNTA FNTB ICMT PCYOX1 RCE1 ZMPSTE24 M_116 (E,E)-Farnesyl-PP_Ubiquinone CLK1 COQ2 COQ3 COQ5 COQ6 COQ7 PDSS1 PDSS2 PHB PPT1 M_117 (E,E)-Farnesyl-PP_Dolichyl phosphate ALG11 DHDDS DOLK SRD5A1 SRD5A2 SRD5A3 M_118 Dolichyl phosphate_Dolichyl phosphate D-mannose DPM1 M_119 Dolichyl phosphate_(GlcNAc)4 (Man)3 (Asn)1 ALG10 ALG10B ALG5 ALG6 ALG8 GANAB MAN1A1 MAN1A2 MAN1B1 MAN1C1 MAN2A1 MAN2A2 MGAT1 MGAT2 MOGS STT3A STT3B M_120 (Glc)3 (GlcNAc)2 (Man)9 (PP-Dol)1+Protein asparagine_(Glc)3 (GlcNAc)2 (Man)9 (Asn)1 STT3A STT3B M_121 (Glc)3 (GlcNAc)2 (Man)9 (Asn)1_(GlcNAc)4 (Man)3 (Asn)1 GANAB MAN1A1 MAN1A2 MAN1B1 MAN1C1 MAN2A1 MAN2A2 MGAT1 MGAT2 MOGS M_122 (GlcNAc)4 (Man)3 (Asn)1_(Gal)2 (GlcNAc)4 (LFuc)1 (Man)3 (Neu5Ac)2 (Asn)1 ABO B4GALT1 B4GALT2 B4GALT3 FUT8 ST6GAL1 ST6GAL2 M_123 (GlcNAc)4 (Man)3 (Asn)1_(GlcNAc)7 (Man)3 (Asn)1 MGAT4A MGAT4B MGAT4C M_124 (GlcNAc)4 (Man)3 (Asn)1_(GlcNAc)5 (Man)3 (Asn)1 MGAT3 M_125 Dolichyl phosphate D-mannose+Protein serine_(Gal)1 (GlcNAc)1 (Man)1 (Ser/Thr)1 ABO B3GLCT B4GALT1 B4GALT2 B4GALT3 FUT4 FUT7 MGAT5B POGLUT1 POMGNT1 POMT1 POMT2 M_126 (Gal)1 (GlcNAc)1 (Man)1 (Ser/Thr)1_(Gal)1 (GlcNAc)1 (Man)1 (Neu5Ac)1 (Ser/Thr)1 ST3GAL2 ST3GAL3 ST3GAL6 M_127 (Gal)1 (GlcNAc)1 (Man)1 (Ser/Thr)1_(Gal)1 (GlcA)1 (GlcNAc)1 (Man)1 (S)1 (Ser/Thr)1 B3GAT2 CHST10 M_128 (Gal)1 (GlcNAc)1 (Man)1 (Ser/Thr)1_(Gal)1 (GlcNAc)1 (LFuc)1 (Man)1 (Ser/Thr)1 FUT9 M_129 Protein serine_(Gal)2 (GlcA)1 (Xyl)1 (Ser)1 B3GALT6 B3GAT3 B4GALT7 XYLT1 XYLT2 M_130 (Gal)2 (GlcA)1 (Xyl)1 (Ser)1_Chondroitin CHPF CHPF2 CHSY1 CHSY3 CSGALNACT1 CSGALNACT2 DSE M_131 Chondroitin_Dermatan DSE M_132 (Gal)2 (GlcA)1 (Xyl)1 (Ser)1_(Gal)2 (GlcA)2 (GlcNAc)2 (Xyl)1 (Ser)1 EXT1 EXT2 EXTL1 EXTL2 EXTL3 M_133 PRPP+Glutamine_IMP ADSL ATIC CANT1 ENTPD1 ENTPD3 ENTPD4 ENTPD5 ENTPD6 ENTPD8 GART NME1 NME1-NME2 NME2 NME3 NME4 NME5 NME6 NME7 NUDT16 NUDT5 PAICS PFAS PPAT M_134 5-Phosphoribosylamine_AICAR ADSL GART PAICS PFAS M_135 AICAR_IMP ATIC CANT1 ENTPD1 ENTPD3 ENTPD4 ENTPD5 ENTPD6 ENTPD8 NME1 NME1-NME2 NME2 NME3 NME4 NME5 NME6 NME7 NUDT16 NUDT5 M_136 IMP_AMP ADSL ADSS ADSSL1 AMPD1 AMPD2 AMPD3 M_137 IMP_XMP IMPDH1 IMPDH2 M_138 AMP_Deoxyadenosine AK1 AK2 AK3 AK4 AK5 AK6 AK7 AK8 ENTPD1 ENTPD3 ENTPD8 F9 NME1 NME1-NME2 NME2 NME3 NME4 NME5 NME6 NME7 NT5C NT5C1A NT5C1B NT5C1B-RDH14 NT5C2 NT5C3A NT5C3B NT5E NT5M PKLR PKM RRM1 RRM2 RRM2B M_139 AMP_Adenine ADK DCK NT5C NT5C1A NT5C1B NT5C1B-RDH14 NT5C2 NT5C3A NT5C3B NT5E NT5M PNP M_140 ADP_Deoxyadenosine AK1 AK2 AK4 AK5 AK7 AK8 NME1 NME1-NME2 NME2 NME3 NME4 NME5 NME6 NME7 NT5C NT5C1A NT5C1B NT5C1B-RDH14 NT5C2 NT5C3A NT5C3B NT5E NT5M PKLR PKM RRM1 RRM2 RRM2B M_141 Deoxyadenosine_Adenine PNP M_142 Deoxyadenosine_Hypoxanthine ADA PNP M_143 IMP_Hypoxanthine ADA HPRT1 NT5C NT5C1A NT5C1B NT5C1B-RDH14 NT5C2 NT5C3A NT5C3B NT5E NT5M PNP M_144 Hypoxanthine_Xanthine XDH M_145 XMP_Xanthine HPRT1 NT5C NT5C1A NT5C1B NT5C1B-RDH14 NT5C2 NT5C3A NT5C3B NT5E NT5M NUDT2 NUDT5 PNP M_146 Xanthine_Uric acid XDH M_147 Xanthine_Guanine GDA M_148 XMP_Guanine APRT GMPS HPRT1 NT5C NT5C1A NT5C1B NT5C1B-RDH14 NT5C2 NT5C3A NT5C3B NT5E NT5M NUDT2 NUDT5 PNP M_149 GMP_Guanine APRT HPRT1 NT5C NT5C1A NT5C1B NT5C1B-RDH14 NT5C2 NT5C3A NT5C3B NT5E NT5M PNP M_150 PRPP_UMP UMPS M_151 Orotidylic acid_UMP UMPS M_152 UMP_B-Alanine DPYD DPYS NT5C NT5C1A NT5C1B NT5C1B-RDH14 NT5C2 NT5C3A NT5C3B NT5E NT5M UCK1 UCK2 UCKL1 UPB1 UPP1 UPP2 UPRT M_153 UMP_CDP CANT1 CMPK1 CMPK2 CTPS1 CTPS2 ENTPD1 ENTPD3 ENTPD4 ENTPD5 ENTPD6 ENTPD8 NME1 NME1-NME2 NME2 NME3 NME4 NME5 NME6 NME7 M_154 Uracil_B-Alanine DPYD DPYS UPB1 M_155 UTP_CDP CTPS1 CTPS2 ENTPD1 ENTPD3 ENTPD8 NME1 NME1-NME2 NME2 NME3 NME4 NME5 NME6 NME7 M_156 CDP_Cytidine CMPK1 CMPK2 ENTPD1 ENTPD3 ENTPD8 NT5C NT5C1A NT5C1B NT5C1B-RDH14 NT5C2 NT5C3A NT5C3B NT5E NT5M UCK1 UCK2 UCKL1 M_157 CDP_dCDP RRM1 RRM2 RRM2B M_158 dCDP_Deoxycytidine CMPK1 CMPK2 DCK DCTPP1 NT5C NT5C1A NT5C1B NT5C1B-RDH14 NT5C2 NT5C3A NT5C3B NT5E NT5M M_159 dCMP_Deoxycytidine DCK NT5C NT5C1A NT5C1B NT5C1B-RDH14 NT5C2 NT5C3A NT5C3B NT5E NT5M M_160 dCDP_dUMP CMPK1 CMPK2 DCTD DCTPP1 M_161 dCDP_dCTP NME1 NME1-NME2 NME2 NME3 NME4 NME5 NME6 NME7 M_162 dUMP_Deoxyuridine NT5C NT5C1A NT5C1B NT5C1B-RDH14 NT5C2 NT5C3A NT5C3B NT5E NT5M TK1 TK2 M_163 dUMP_dTMP TYMS M_164 dTMP_Succinyl-CoA ACSF3 AGXT2 ALDH1B1 ALDH2 ALDH3A2 ALDH7A1 ALDH9A1 AOX1 DPYD DPYS MUT NT5C NT5C1A NT5C1B NT5C1B-RDH14 NT5C2 NT5C3A NT5C3B NT5E NT5M TK1 TK2 TYMP UPB1 M_165 dTMP_dTTP DTYMK ENTPD1 ENTPD3 ENTPD8 NME1 NME1-NME2 NME2 NME3 NME4 NME5 NME6 NME7 M_166 Thymine_Succinyl-CoA ACSF3 AGXT2 ALDH1B1 ALDH2 ALDH3A2 ALDH7A1 ALDH9A1 AOX1 DPYD DPYS MUT UPB1 M_167 (E,E)-Farnesyl-PP_Cholesterol CYP51A1 DHCR24 DHCR7 EBP FDFT1 HSD17B1 HSD17B12 HSD17B2 HSD17B6 HSD17B7 HSD17B8 LSS MSMO1 NSDHL RNF2 SC5D SQLE TM7SF2 M_168 Cholesterol_Chenodeoxycholate ACOT1 ACOT2 ACOT4 ACOT7 ACOT8 ACOX2 AKR1C4 AKR1D1 AMACR BAAT CH25H CYP27A1 CYP39A1 CYP46A1 CYP7A1 CYP7B1 CYP8B1 HSD11B2 HSD17B4 HSD3B7 RDH10 RDH16 RDH8 SCP2 SLC27A5 M_169 Cholesterol_ Steroid_hormone AKR1C1 AKR1C2 AKR1C3 AKR1C4 AKR1D1 ALG11 COMT CYP11A1 CYP11B1 CYP11B2 CYP17A1 CYP19A1 CYP1A1 CYP1A2 CYP1B1 CYP2A13 CYP2A6 CYP2A7 CYP2B6 CYP2C18 CYP2C19 CYP2C8 CYP2C9 CYP2D6 CYP2E1 CYP2F1 CYP2J2 CYP2S1 CYP3A4 CYP3A43 CYP3A5 CYP3A7 CYP4B1 CYP4F8 CYP4X1 CYP4Z1 HSD11B1 HSD11B2 HSD17B1 HSD17B12 HSD17B2 HSD17B3 HSD17B6 HSD17B7 HSD17B8 HSD3B1 HSD3B2 RDH10 RDH16 RDH8 RNF2 SRD5A1 SRD5A2 SRD5A3 STS SULT1E1 SULT2B1 UGT1A1 UGT1A10 UGT1A3 UGT1A4 UGT1A5 UGT1A6 UGT1A7 UGT1A8 UGT1A9 UGT2A1 UGT2A2 UGT2A3 UGT2B10 UGT2B11 UGT2B15 UGT2B17 UGT2B28 UGT2B4 UGT2B7 M_170 XMP_GMP GMPS M_171 dCDP_dCMP CMPK1 CMPK2 DCTPP1